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Before the Roman invasion, Britain was populated by Celtic tribes with well-established cultural and economic links with continental Europe. Although Julius Caesar conducted the first Roman campaign in Britain in 55 B.C., the conquest did not begin until A.D. 43, during the reign of Claudius. The British tribes initially opposed the Roman legions, but by 84 the Romans had decisively conquered southern Britain and had pushed into what is now southern Scotland. In 122 they fortified the northern border with Hadrian's Wall, which spanned what is now Northern England. In 142 Roman forces pushed north again and began construction of the Antonine Wall, but they retreated back to Hadrian's Wall after only twenty years. Following the conquest, native Britons were subject to the Roman governors but mostly kept their land, and a distinctive Romano-British culture emerged. The Roman Empire retained control until its departure about A.D. 430.